A wealthy American dentist who used a bow and arrow to shoot Zimbabwe's beloved Cecil the lion faced fierce criticism Tuesday amid allegations that it took 40 hours for the animal to die.
The lion, a popular attraction among international visitors to the Hwange National Park, was lured outside the park's boundaries by bait and killed earlier this month.
"The offender is a gentleman who is an American called Walter James Palmer," Emmanuel Fundira, president of the Safari Operators Association of Zimbabwe, told reporters in the capital Harare.
"A lot of people travel long distances coming to Zimbabwe to enjoy our wildlife and obviously the absence of Cecil is a disaster."
Fundira described Cecil as "almost semi-domesticated" and said that Palmer had paid about $50,000 for the hunt.
The Zimbabwe Conservation Task Force charity said Palmer and professional Zimbabwean hunter Theo Bronkhorst had gone out at night with a spotlight and tied a dead animal to their vehicle to lure Cecil into range.
"Palmer shot Cecil with a bow and arrow but this shot didn't kill him. They tracked him down and found him 40 hours later when they shot him with a gun," the charity alleged.
It added that the hunters unsuccessfully tried to hide the dead lion's tracking collar, which was part of a University of Oxford research programme.
"Cecil was skinned and beheaded. We don't know the whereabouts of the head," the charity alleged.
Cecil, aged about 13, was said by safari operators to be an "iconic" animal who was recognised by many visitors to Hwange due to his distinctive black mane.
Hwange attracted 50,000 visitors last year, about half of them from abroad.
Bronkhorst and local landowner Honest Ndlovu will appear in court in Victoria Falls on Wednesday to face poaching charges.
"Both the professional hunter and land owner had no permit or quota to justify the offtake of the lion and therefore are liable for the illegal hunt," the Zimbabwean Parks Authority said in a statement Tuesday.
"(They) are being jointly charged for illegally hunting the lion."
The statement made no mention of Palmer, but added that Bronkhorst's son Zane was also wanted for questioning.
Palmer is an experienced trophy hunter from Minnesota.
He is well-known in US hunting circles as an expert shot with his bow and arrow, and has travelled across the world in pursuit of leopards, buffalo, rhino, elk and other large mammals.
Palmer, who reportedly pleaded guilty in 2008 to poaching a black bear in Wisconsin, said he had "relied on the expertise of my local professional guides to ensure a legal hunt."
"I deeply regret that my pursuit of an activity I love and practise responsibly and legally resulted in the taking of this lion," Palmer said in a statement.
"I had no idea that the lion I took was a known, local favourite, was collared and part of a study until the end of the hunt."
The Twitter and Facebook accounts and website of his dental practice - River Bluff Dental in a suburb of Minneapolis, Minnesota - were shut down Tuesday after being flooded with blistering attacks.
A makeshift memorial began forming outside his shuttered office as people outraged by the story dropped off stuffed animals and flowers, according to images from local media.
The online outrage was intense and by late afternoon, there were nearly 100,000 tweets with the hashtag #CecilTheLion, while an online petition demanding justice for Cecil had drawn 95,000 signatures.
Comedian Ricky Gervais, who has targeted trophy hunters with his barbs, was among those who expressed their disgust.
"It's not for food. It's not the shooting, or tin cans would do. It must just be the thrill of killing. Mental," Gervais tweeted.
Yelp reviewer Ana S. wrote: "You are a worthless human being."
"Isn't that the guy who butchered all those animals for some psychotic thrill?" a commentator identified as OldBoy wrote on his YouTube page.
Some hunting of lions and other large animals is legal in countries such as South Africa, Namibia and Zimbabwe, where permits are issued allowing hunters to kill certain beasts that are beyond breeding age.
It is outlawed in Zimbabwe's national parks.
"We use hunting as a conservation tool," Fundira said. "Hunting is only based on animals which have been vigorously checked in terms of qualifying."
The Zimbabwe Conservation Task Force said Cecil had cubs, which were likely to be killed by the lion that took over in the group's hierarchy.
In May, a Texan hunter shot an elderly black rhinoceros in Namibia after buying a government-issued licence for $350,000.